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The cerebellum, principally a motor organ, is responsible for the coordination of movements, especially skilled voluntary ones; the control of posture and gait, and the regulation of muscular tone The mechanisms by which these functions are accomplished have been the subject of intense investigation by anatomists and physiologists Their studies have yielded a mass of data, testimony to the complexity of the organization of the cerebellum and its afferent and efferent connections A coherent picture of cerebellar function is now emerging, although it is not yet possible, with a few notable exceptions, to relate each of the symptoms of cerebellar disease to a derangement of a discrete anatomic or functional unit of the cerebellum Knowledge of cerebellar function has been derived mainly from the study of natural and experimental ablative lesions and to a lesser extent from stimulation of the cerebellum, which produces little in the way of movement or alterations of induced movement Notably, none of the motor activities of the cerebellum reach conscious kinesthetic perception; its main role is to assist in the initiation and modulation of willed movements that are generated in the cerebral hemispheres The following outline of cerebellar structure and function has of necessity been simpli ed; a full account can be found in the writings of Jansen and Brodal, Gilman, and Thach and colleagues, listed at the end of this chapter

The classic studies of the comparative anatomy and ber connections of the cerebellum have led to its subdivision into three parts (Fig 5-1): (1) The occulonodular lobe, located inferiorly, which is phylogenetically Central lobule the oldest portion of the cerebellum and is Lingula Anterior lobe much the same in all animals (hence archi(Paleocerebellum or Primary fissure cerebellum) It is separated from the main Spinocerebellum) t g An dran e Posterior superior mass of the cerebellum, or corpus cerebelli, a ul fissure Qu Lob Si by the posterolateral ssure (2) The anterior Culmen Lo mp ar bu le st gul lobe, or paleocerebellum, which is the porle Po ran le Su d u tion of the corpus cerebelli rostral to the priSemperior ua Lob Declive Q Lob ilunar mary ssure In lower animals, the anterior CRUS I ule Horizontal Folium lobe constitutes most of the cerebellum, but fissure Tuber in humans it is relatively small, consisting of ior r r Posterior lobe Pyramis Infe iluna CRUS II the anterosuperior vermis and the contiguous (Neocerebellum or le Sem bule bu pontocerebellum) paravermian cortex (3) The posterior lobe, Lo lo Uvula or neocerebellum, consisting of the middle Prepyramidal divisions of the vermis and their large lateral fissure extensions The major portions of the human Floccuodular lobe Posterolateral fissure cerebellar hemispheres fall into this, the larg(Archicerebellum or Nodulus Flocculus Vestibulocerebellum) est, subdivision This anatomic subdivision corresponds Paraflocculus roughly with the distribution of cerebellar Figure 5-1 Diagram of the cerebellum, illustrating the major ssures, lobes, and lobules and the function, based on the arrangement of its af- major phylogenetic divisions (left)

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